(1) National Institute of Agronomy of Tunisia (INAT), Research Unit UR17AGR01 “Valorization of the Tunisian natural and agro-food heritage through innovation”, University of Carthage, 43 Avenue Charles Nicolle, 1082 Tunis, Tunisia ,

(2) Food Science Research Institute, National Agricultural Research and Innovation Centre, Herman Otto utca 15, H-1022 Budapest, Hungary ,

(3) Food Science Research Institute, National Agricultural Research and Innovation Centre, Herman Otto utca 15, H-1022 Budapest, Hungary ,

(4) Food Science Research Institute, National Agricultural Research and Innovation Centre, Herman Otto utca 15, H-1022 Budapest, Hungary ,

(5) Food Science Research Institute, National Agricultural Research and Innovation Centre, Herman Otto utca 15, H-1022 Budapest, Hungary ,

(6) Food Science Research Institute, National Agricultural Research and Innovation Centre, Herman Otto utca 15, H-1022 Budapest, Hungary ,

(7) National Institute of Cereal crops (INGC), 8170 Bou Salem, Tunisia ,

(8) National Institute of Agronomy of Tunisia (INAT), Genetic and Plant Breeding Laboratory, University of Carthage, 43 Avenue Charles Nicolle, 1082 Tunis, Tunisia ,

(9) National Institute of Agronomy of Tunisia (INAT), Genetic and Plant Breeding Laboratory, University of Carthage, 43 Avenue Charles Nicolle, 1082 Tunis, Tunisia

(1) Laboratoire de Nutrition, Pathologie, Agro-biotechnologie et Santé (Lab-NuPABS). Department of Biology, Faculty of Natural and Life Sciences, Djillali Liabes University, Sidi-Bel-Abbes, Algeria ,

(2) National Institute of Agronomy of Tunisia (INAT), Department of Agri-food Industries, University of Carthage, 43 Avenue Charles Nicolle, 1082 Tunis, Tunisia ,

(3) Département des Sciences Biologiques, Faculté des Sciences de la Nature et de la Vie et des Sciences de la Terre, Université de Bouira, 10000, Bouira, Algeria ,

(4) Research Scientist Joint Research Unit in Nutrition and Food Regional Designated Center of Nutrition AFRA/IAEA, (Ibn Tofail University-CNESTEN), Morocco ,

(5) London School of Hygiene & Tropical Medicine, Faculty of Public Health and Policy, 15-17 Tavistock Place, London WC1H 9SH, UK, United Kingdom

(1) Laboratory of cereal technology (LR20INRAT02), National Institute of Agronomic Research of Tunis, Carthage University, Hédi Karray street – 2049 Ariana -Tunisia. National Agronomic Institute of Tunisia (UR17AGR01), Carthage University, 43, Charles Nicolle Avenue – 1082 Tunis- Mahrajène, Tunisia ,

(2) Laboratory of cereal technology (LR20INRAT02), National Institute of Agronomic Research of Tunis, Carthage University, Hédi Karray street – 2049 Ariana, Tunisia ,

(3) Competitiveness pole of Bizerte,Union du Grand Maghreb Arabe boulevard - 7080 Menzel Jemil, Bizerte, Tunisia ,

(4) National Agronomic Institute of Tunisia (UR17AGR01), Carthage University, 43, Charles Nicolle Avenue – 1082 Tunis- Mahrajène, Tunisia

(1) Department of Agri-Food Industries, National Institute of Agronomy of Tunisia (INAT), UR17AGR01, University of Carthage, Tunis. Faculty of Mathematical, Physical and Natural Sciences of Tunis, LR11ES09, University Tunis El Manar, Tunis, Tunisia ,

(2) Department of Agri-Food Industries, National Institute of Agronomy of Tunisia (INAT), UR17AGR01, University of Carthage, Tunis, Tunisia ,

(3) National Institute for Consumption (INC), Tunis, Tunisia ,

(4) Department of Agri-Food Industries, National Institute of Agronomy of Tunisia (INAT), UR17AGR01, University of Carthage, Tunis, Tunisia

(1) University of Carthage, National Institute of Agronomy of Tunisia (INAT), Department of Agri-Food Industries, UR17AGR01, 1082 Tunis, Tunisia ,

(2) University of Jendouba, Institut Superieur de Biotechnologie de Beja, Avenue Habib Bourguiba, 9000 Béja, Tunisia ,

(3) University of Carthage, National Institute of Agronomy of Tunisia (INAT), Department of Agri-Food Industries, UR17AGR01, 1082 Tunis, Tunisia