(1) Regional Center of Agricultural Research of Sidi Bouzid (CRRA) PB 357, 9100 Sidi Bouzid,, Tunisia ,

(2) Regional Center for Agricultural Research (CRRA), B.P 357, Sidi Bouzid, 9100,, Tunisia ,

(3) Institut Supérieur des Etudes Technologiques de Sidi Bouzid, 9100,, Tunisia ,

(4) Institut Supérieur des Etudes Technologiques de Sidi Bouzid, 9100,, Tunisia ,

(5) Institut Supérieur des Etudes Technologiques de Sidi Bouzid, 9100,, Tunisia

(1) Laboratoire BIOQUAL, Département de Biotechnologie, INATAA, UFMC1, Algeria ,

(2) Laboratoire BIOQUAL, Département de Biotechnologie, INATAA, UFMC1, Algeria

(1) Department of Microbiology and Biochemistry, Faculty of Life and Natural Science, University of Batna2, Route de Constantine, 05078, Fesdis, Batna, Algeria ,

(2) Department of Microbiology and Biochemistry, Faculty of Life and Natural Science, University of Batna2, Route de Constantine, 05078, Fesdis, Batna, Algeria