(1) Moulay Ismail University Faculty of Sciences Meknes Laboratory of Biotechnology and Bioresources Valorization BP 11201, Zitoune, Meknes, Morocco ,

(2) Moulay Ismail University Cluster of Competency «Agri-food, Safety and Security» IUC VLIR-UOS Marjane 2, BP 298, Meknes, Morocco